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Covering 100 years of zoological research the handbook of zoology online represents a vast store of knowledge the handbook of zoology offers an in depth treatment of the entire animal kingdom covering both invertebrates and vertebrates internationally renowned experts provide comprehensive overviews on animal systematics and morphology as well as thorough discussions of physiology . Handbook of zoology handbuch der zoologie handbook of zoology arthropoda insecta coleoptera handbook of zoology handbuch der zoologie handbook and . Handbuch der zoologie handbook of zoology vol iv arthropoda insecta part 38 coleoptera beetles volume 1 morphology and systematics edited by rg beutel rab leschen walter de gruyter berlin and new york 2005 xi 567 pp isbn 3 11 017130 9 us33480. Handbuch der zoologie handbook of zoology vol iv arthropoda insecta part 38 coleoptera beetles volume 1 morphology and systematics edited by rg beutel rab leschen. This book is a revised edition of the first of three volumes in the handbook of zoology series which treats the systematics and biology of coleoptera with over 380000 described species coleoptera are by far the most species rich order of insects and the largest group of animals of comparable geological age moreover numerous species are tremendously important economically